Richard E. Clopton, Ph.D.

Curriculum Vita


The Downey Family Honors Chair in Science
Professor of Biology
Department of Natural Science
Peru State College, Peru, NE 68421 USA
402/872-2284
rclopton@peru.edu

EDUCATION

1993-1995: Postdoctoral Research Associate, Texas A&M University, Dept of Entomology, College Station, Texas
1990-1993: PhD, University of Nebraska, Parasitology, "Specificity in the gregarine assemblage parasitizing Tenebrio molitor.",
1987-1989: MS, University of Nebraska, Entomology, "Some aspects of the biology and ecology of the common pest chigger mite, Eutrombicula alfreddugesi, Oudemans 1912."
1983-1987: BS, University of Nebraska, Entomology

RECENT GRANTS

2010-2014: NSF: Biodiversity Inventory: Gregarines Parasitizing North American Cockroaches.
2010-2012: NSF: Primarily Undergraduate Institutions: An Important Resource for Systematics Research.
2004-2007: NSF: Microbiological Survey & Inventory of gregarines parasitizing aquatic and riparian insects of the Texas Big Thicket.
1997-2001: NSF: Microbiological Survey & Inventory: Eugregarinida (Protista: Apicomplexa) parasitizing selected mandibulate arthropods of the Nebraska Sandhills.

PUBLICATIONS

Steele, S. M., D. T. Clopton, and R. E. Clopton. 2012. Excystation signals do not isolate gregarine gene pools: experimental excystation of Blabericola migrator among eleven species of cockroaches. Journal of Parasitology 98(5): 946-950.PDF

Clopton, R. E. 2012. Redescription of Protomagalhaensia blaberae Peregrine, 1970 (Apicomplexa: Eugregarinida: Blabericolidae) parasitizing the Bolivian Cockroach, Blaberus boliviensis (Dictyoptera: Blaberidae). Comparative Parasitology 79(2): 181-191. PDF

Trout, K. and R. E. Clopton. 2012. Metronidazole induces gametocytogenesis in gregarine associations maintained in vitro. Journal of Parasitology 98(3):666-668. PDF

Clopton, R. E. 2012. Synoptic revision of Blabericola (Apicomplexa: Eugregarinida: Blabericolidae) parasitizing blaberid cockroaches (Dictyoptera: Blaberidae) with comments on recognizing and delineating gregarine species boundaries. Journal of Parasitology 98(3): 572-583. PDF

Clopton, R. E. 2011. Redescription of Protomagalhaensia granulosae Peregrine, 1970 (Apicomplexa: Eugregarinida: Blabericolidae) parasitizing the Discoid Cockroach, Blaberus discoidalis (Dictyoptera: Blaberidae). Comparative Parasitology 78(1):63-72. PDF

Cielocha, J. J., T. J. Cook, R. E. Clopton. 2011. Host Utilization and Distribution of Nubenocephalid Gregarines (Eugregarinorida: Actinocephalidae) Parasitizing Argia spp. (Odonata: Zygoptera) in the Central United States. Comparative Parasitology 78(1):152-160. PDF

Cook, T. J., R. E. Clopton, And J. L. Cook. 2011. Neoasterophora megasylvaticus n. gen., n. sp. (Apicomplexa: Eugregarinida: Actinocephalidae: Pileocephalinae) Parasitizing Cheumatopsyche analis (Trichoptera: Hydropsychidae) in the Texas Big Thicket. Comparative Parasitology 78(1):56-62. PDF

Clopton, R. E., T. J. Cook, and J. J. Cielocha. 2010. Nubenocephalus nickoli n. sp. and Nubenocephalus xunantunichensis n. sp. (Apicomplexa: Eugregarinida: Actinocephalidae) Parasitizing Damselflies (Odonata: Zygoptera) in Belize, Central America. Comparative Parasitology 77(2):125-136. PDF

Clopton, R. E. 2010. Protomagalhaensia cerastes n. sp. (Apicomplexa: Eugregarinida: Blabericolidae) Parasitizing the Pallid Cockroach, Phoetalia pallida (Dictyoptera: Blaberidae). Comparative Parasitology 77(2): 117-124. PDF

Clopton, R. E. 2009. Phylogenetic Relationships, Evolution, and Systematic Revision of the Septate Gregarines (Apicomplexa: Eugregarinorida: Septatorina). Comparative Parasitology 76(2): 167-190. PDF

Clopton, R. E., T. J. Cook, and J. L. Cook. 2008. Gregarina tropica n. sp. (Apicomplexa: Eugregarinorida: Gregarinicae: Gregarinidae) Parasitizing the Brown-winged Earwig, Vostox brunneipennis (Dermaptera: Labiidae), in the Texas Big Thicket. Comparative Parasitology 75:215-227. PDF

Clopton, R. E., T. J. Cook, and J. L. Cook. 2008. Trichurispora wellgundis n. g., n. sp. (Apicomplexa: Eugregarinida: Hirmocystidae) parasitizing adult Water Scavenger Beetles, Tropisternus collaris (Coleoptera: Hydrophilidae) in the Texas Big Thicket. Comparative Parasitology 75: 82-91. PDF

Realzola, E., J. L. Cook, T. J. Cook, and R. E. Clopton. 2007. Composition of gyrinid aggregations in the East Texas Primitive Big Thicket. Coleopterist's Bulletin 61:495-502.

Hays, J. J., R. E. Clopton, T. J. Cook and J. L. Cook. 2007. Revision of the Genus Nubenocephalus and Description of Nubenocephalus secundus N. Sp. (Apicomplexa: Actinocephalidae) Parasitizing Adults of Argia sedula (Odonata: Zygoptera: Coenagrionidae) in the Primitive Texas Big Thicket, U.S.A. Comparative Parasitology. 74: 286-293. PDF

Clopton, R.E., T. J. Cook and J. L. Cook. 2007. Revision of Geneiorhynchus Schneider, 1875 (Apicomplexa: Eugregarinida: Actinocephalidae: Acanthosporinae) with Recognition of 4 New Species of Geneiorhynchus and Description of Geneiorhynchus manifestus N. Sp. Parasitizing Naiads of the Green Darner, Anax junius (Odonata: Aeshnidae) in the Texas Big Thicket. Comparative Parasitology 74: 273-285. PDF

Clopton, R. E., and J. J. Hays. 2006. Revision of the Genus Protomagalhaensia and Description of Protomagalhaensia wolfi n. comb. (Apicomplexa: Eugregarinida: Hirmocystidae) and Leidyana haasi n. comb. (Apicomplexa: Eugregarinida: Leidyanidae) Parasitizing the Lobster Cockroach, Nauphoeta cinerea (Dictyoptera: Blaberidae) Comparative Parasitology 73: 137-156. PDF

Clopton, R. E. 2006. Two New Species of Xiphocephlaus in Eleodes tricostata and Eleodes fusiformis (Coleoptera: Tenebrionidae: Eleodini) from The Sandhills of Western Nebraska. Journal of Parasitology 92: 569-577. PDF

Welsh, D., R. E. Clopton, and L. Parris. 2006. Differential Temperature Acclimatization Responses in the Membrane Phospholipids of Posthodiplostomum minimum and its Second Intermediate Host, Lepomis macrochirus. Journal of Parasitology 92: 764-769. PDF

Snyder, S. D., and R. E. Clopton. 2005. New Methods for the Collection and Preservation of Spirorchiid and Polystomatid Trematodes from Turtles. Comparative Parasitology 72: 102-107. PDF

Hays, J., R. E. Clopton, D. L. Cappaert, and D. R. Smitley. 2004. Revision of the genus Stictospora and description of Stictospora villani, n. sp. (Apicomplexa: Eugregarinida: Actinocephalidae) from the Japanese beetle Popillia japonica (Coleoptera: Scarabaeidae) in Michigan, USA. Journal of Parasitology 90: 1450-1456. PDF

Clopton, R. E. 2004. Standard nomenclature and metrics of plane shapes for use in gregarine taxonomy. Comparative Parasitology 71: 130-140. PDF

Clopton, R. E. 2004. Calyxocephalus karyopera g. nov., sp. nov. (Eugregarinorida: Actinocephalidae: Actinocephalinae) from the ebony jewelwing damselfly Calopteryx maculata (Zygoptera: Calopterygidae) in southeast Nebraska: implications for mechanical prey-vector stabilization of exogenous gregarine development. Comparative Parasitology 71: 141-153. PDF

Clopton, R. E., T. J. Cook, and J. L. Cook. 2004. Naiadocystis phykoterion n. gen., n. sp. (Apicomplexa: Eugregarinida: Hirmocystidae) from the mexican pygmy grasshopper Paratettix mexicanus (Orthoptera: Tetrigidae) in the Texas Big Thicket with recognition of three previously described species of Naiadocystis. Journal of Parasitology 90: 301-307. PDF

Clopton, R. E. 2004. Comparative Parasitology policies and instructions for authors. Comparative Parasitology 70: 218-225.

Smith A., and R. E. Clopton. 2003. Efficacy of oral metronidazole and potassium sorbate against two gregarine parasites, Protomagalhaensia granulosae and Gregarina cubensis (Apicomplexa: Eugregarinida) infecting the Death's Head Cockroach, Blaberus discoidalis. Comparative Parasitology 70: 196-199. PDF

Leander, B. S., R. E. Clopton, and P. J. Keeling. 2003. Phylogeny of gregarines (Apicomplexa) as inferred from small-subunit rDNA and beta-tubulin. International Journal of Systematic and Evolutionary Microbiology 53: 345-354. PDF

Clopton, R. E., and C. M. Nolte. 2002. Clitellocephalus americanus n. gen., n. sp. (Apicomplexa: Eugregarinida: Gregarinidae) from Cratacanthus dubius (Coleoptera: Carabidae: Harpalinae) in the Nebraska Sandhills and Clitellocephalus ophoni n. comb. (Apicomplexa: Eugregarinida: Gregarinidae) from Ophonus pubescens (Coleoptera: Carabidae: Harpalinae) in Sète, France. Journal of Parasitology 88: 750-757. PDF

Clopton, R. E., and A. Smith. 2002. Efficacy of oral sulfadimethoxine against two gregarine parasites, Protomagalhaensia granulosae and Gregarina cubensis (Apicomplexa: Eugregarinida) infecting the Death's Head Cockroach, Blaberus discoidalis. Journal of Parasitology 88: 786-789. PDF

Clopton, R. E. 2002. Phylum Apicomplexa Levine, 1970: Order Eugregarinorida Léger, 1900. Pages 205-288 in: Illustrated Guide to the Protozoa, 2nd edition, J. J. Lee, G. Leedale, D. Patterson, and P. C. Bradbury, eds. Society of Protozoologists, Lawrence, Kansas. PDF

Cook, T. J., J. Janovy, Jr., and R. E. Clopton. 2001. Epimerite-host epithelium relationships among eugregarines parasitizing the damselflies Enallagma civile and Ischnura verticalis. Journal of Parasitology 87: 988-996. PDF

Clopton, R. E. 2000. Stylocephalus occidentalis n. sp.(Apicomplexa: Eugregarinida: Stylocephalidae) from Trimytis pruinosa (Coleoptera: Tenebrionidae) in the Nebraska Sandhills. Journal of Parasitology 86: 560-565. PDF

Kula, R. R. and R. E. Clopton. 1999. Amoebogregarina nigra n. gen., n. comb. (Apicomplexa: Gregarinidae) From Adult Melanoplus differentialis (Orthoptera: Acrididae) in Southeastern Nebraska. Journal of Parasitology 85: 321-325. PDF

Clopton, R. E. 1999. Revision of the genus Xiphocephalus and description of Xiphocephalus ellisi n. sp.(Apicomplexa: Eugregarinida: Stylocephalidae) from Eleodes opacus (Coleoptera: Tenebrionidae) in the western Nebraska Sandhills. Journal of Parasitology 85. 84-89. PDF

Lucarotti, C. J., T. L. Leclerc, and Clopton, R. E. 1998. Incidence of Leidyana canadensis (Apicomplexa: Eugregarinida) in Lambdina fiscellaria fiscellaria larvae (Lepidoptoera: Geometridae). Canadian Entomologist 130: 583-594. PDF

Clopton, R. E. 1998. Revision of the genus Torogregarina and description of Torogregarina sphinx n. sp. (Apicomplexa: Eugregarinida) from a Missouri river bank beetle, Bembidion laevigatum (Coleoptera: Carabidae), in southeastern Nebraska. Journal of Parasitology 84: 823-827. PDF

Clopton, R. E. and C. J. Lucarotti. 1997. Leidyana canadensis N. Sp. (Apicomplexa: Eugregarinida) from larval Eastern Hemlock Looper, Lambdina fiscellaria fiscellaria (Lepidoptoera: Geometridae). Journal of Eukaryotic Microbiology 44: 383-387. PDF

Janovy, Jr., S. D. Snyder, and R. E. Clopton. 1997. Evolutionary constraints on population structure: the parasites of Fundulus zebrinus (Pisces: Cyrinodontidae) in the South Platte River of Nebraska. Journal of Parasitology 83: 584-592. PDF

Clopton, R. E. and R. E. Gold. 1996. Host Specificity of Gregarina blattarum among Five Species of Domiciliary Cockroaches. Journal of Invertebrate Pathology 67: 219-223.PDF

Clopton, R. E. 1995. Leidyana migrator n. sp. (Apicomplexa: Leidyanidae) from the Madagacar hissing cockroach, Gromphadorhina portentosa (Insecta: Blattodea). Invertebrate Biology 114: 271-278. PDF

Clopton, R. E. 1995. Domadracunculus janovyii n. gen., n. sp., (Apicomplexa: Actinocephalidae) described from adults of Ischnura verticalis (Odonata: Zygoptera) in Texas. Journal of Parasitology 81:256--260. PDF

Clopton, R. E. and R. E. Gold. 1995. Effects of pH on excystation in Gregarina cuneata and Gregarina polymorpha (Eugregarinida: Gregarinidae). Journal of Eukaryotic Microbiology 42: 540-544.PDF

Percival, T. J., R. E. Clopton and J. Janovy, Jr. 1995. Hoplorhynchus acanthatholius N. Sp. and Steganorhynchus dunwoodii N. Gen. N. Sp. (Apicomplexa: Eugregarinorida) described from Coenagrionid Damselflies. Journal of Eukaryotic Microbiology 42: 406-410. PDF

Janovy, Jr., R. E. Clopton, D. A. Clopton, S. Snyder, A. Efting and L. Krebs. 1995. Species density distributions as null models for ecologically significant interactions of parasite species in an assemblage. Ecological Modeling 77: 189-196. PDF

Clopton, R. E. and R. E. Gold. 1993. Distribution and Seasonal and diurnal activity patterns of Eutrombicula alfreddugesi (Acari: Trombiculidae) in a forest edge ecosystem. Journal of Medical Entomology 30: 47--53.

Clopton, R. E. and J. Janovy, Jr. 1993 Developmental niche structure in the gregarine assemblage parasitizing Tenebrio molitor. Journal of Parasitology 79:701--709. PDF

Clopton R. E., T. J. Percival and John Janovy, Jr. 1993. Nubenocephalus nebraskensis N. Gen., N. Sp. (Apicomplexa: Actinocephalidae) described from Argia bipunctulata (Odonata: Zygoptera). Journal of Parasitology 79: 533--537. PDF

Ferdig, M. T., M. A. McDowell, R. E. Clopton and Janovy, J. Jr. 1993. Patterns of morphological variation of Salsuginus yutanensis (Monogenera: Ancyrocephalidae) over space and time. Journal of Parasitology 79: 744-750.

Clopton, R. E. and R. E. Gold. 1992. Bite-Count evaluation of the repellency of N,N-Diethyl-3-methylbenzamide to larval Eutrombicula alfreddugesi (Acari: Trombiculidae). Journal of Medical Entomology 29: 858-863. PDF

Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. Host Stadium Specificity in the gregarine assemblage parasitizing Tenebrio molitor. Journal of Parasitology 78: 334--337. PDF

Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. Gregarina coronata N. Sp. (Apicomplexa: Eugregarinida) described from adults of the Southern Corn Rootworm, Diabrotica undecimpunctata howardi. Journal of Protozoology 39: 417--420. PDF

Janovy, Jr., R. E. Clopton and T. J. Percival. 1992. The Roles of ecological and evolutionary influences in providing structure to parasite species assemblages. Journal of Parasitology 78: 630-640.

Clopton R. E., T. J. Percival and John Janovy, Jr. 1991. Gregarina niphandrodes N. Sp. (Apicomplexa: Eugregarinorida) from adult Tenebrio molitor (L.) with oocyst descriptions of other gregarine parasites of the Yellow Mealworm. Journal of Protozoology 38:472--483. PDF

Clopton R. E. 1991. A review of the Scaritinid Beetles (Coleoptera: Carabidae) of Nebraska. Transactions of the Nebraska Academy of Science 18: 53--65.

home

 

 

 

 

 

Mayan temple complex at Caracol, Cayo District, Belize, 2008.