Richard E. Clopton, PhD
Professor of Biology
Editor, Comparative Parasitology
Downey Family Honors Chair in Science
Department of Natural Science
Peru State College
Peru, NE 68421
402/872-2284
402/872-2255
rclopton@oakmail.peru.edu

Education
1990-1993
PhD, University of Nebraska, Parasitology
Dissertation (under John Janovy, Jr.): "Specificity in the gregarine assemblage parasitizing Tenebrio molitor."
1987-1989
MS, University of Nebraska, Entomology
Thesis (under Roger E. Gold): "Some aspects of the biology and ecology of the common pest chigger mite, Eutrombicula alfreddugesi, Oudemans 1912."
1983-1987
BS, University of Nebraska, Agriculture, Entomology
Grantsmanship
1998
National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 1998-May 15, 1999; $10,000
1997
National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 1997-May 15, 1998; $10,000
Click on the PDF icon to download reprints in PDF format where available.
National Science Foundation.
Microbiological Survey & Inventory: Eugregarinida (Protista: Apicomplexa) parasitizing selected mandibulate arthropods of the Nebraska Sandhills. NSF DEB-9705179.
Funded: May 15, 1997-May 15, 2000; $199,780
Refereed Publications
Professor of Biology
Editor, Comparative Parasitology
Downey Family Endowed Honors Chair in Science
Department of Natural Science
Peru State College, Peru, NE
Lecture responsibilities (1995-Present)
Introductory Biology, Honors Biology, Zoology, Botany, Horticulture, Human Physiology, Genetics, Plant Systematics, Invertebrate Zoology, Entomology, Comparative Vertebrate Anatomy, Human Clinical Parasitology, and Experimental Biology
Scientific research
Systematics & Taxonomy of Gregarines parasitizing mandibulate arthropods; Evolutionary systematics; Evolution of symbiosis
Extramural funding by the National Science Foundation
Strong undergraduate research component
Summer Biological Survey & Inventory Expeditions, 1997 - 2005
1999
National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 1999-May 15, 2000; $10,000
2000
National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 2000-May 15, 2001; $10,000
Position
Richard E. Clopton, PhD
Professor of Biology
Editor, Comparative Parasitology
Downey Famil Honors Chair in Science
Department of Natural Science
Peru State College
Peru, NE 68421
402/872-2284
402/872-2255
rclopton@oakmail.peru.edu

2004
National Science Foundation
Microbiological Survey & Inventory of gregarines parasitizing aquatic and riparian insects of the Texas Big Thicket.
NSF DEB 0340774
Funded: Oct 1. 2004-2007; $200,000

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Snyder, S. D., and R. E. Clopton. 2005. New Methods for the Collection and Preservation of Spirorchiid and Polystomatid Trematodes from Turtles. Comparative Parasitology 72: 102-105. |
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Hays, J., R. E. Clopton, D. L. Cappaert, and D. R. Smitley. 2004. Revision of the genus Stictospora and description of Stictospora villani, n. sp. (Apicomplexa: Eugregarinida: Actinocephalidae) from the Japanese beetle Popillia japonica (Coleoptera: Scarabaeidae) in Michigan, USA. Journal of Parasitology 90:1450-1456. |
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Clopton, R. E. 2004. Standard nomenclature and metrics of plane shapes for use in gregarine taxonomy. Comparative Parasitology 71: 130-140. |
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Clopton, R. E. 2004. Calyxocephalus karyopera g. nov., sp. nov. (Eugregarinorida: Actinocephalidae: Actinocephalinae) from the ebony jewelwing damselfly Calopteryx maculata (Zygoptera: Calopterygidae) in southeast Nebraska: implications for mechanical prey-vector stabilization of exogenous gregarine development. Comparative Parasitology 71: 141-153. |
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Clopton, R. E., T. J. Cook, and J. L. Cook. 2004. Naiadocystis phykoterion n. gen., n. sp. (Apicomplexa: Eugregarinida: Hirmocystidae) from the mexican pygmy grasshopper Paratettix mexicanus (Orthoptera: Tetrigidae) in the Texas Big Thicket with recognition of three previously described species of Naiadocystis. Journal of Parasitology 90: 301-307. |
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Smith A., and R. E. Clopton. 2003. Efficacy of oral metronidazole and potassium sorbate against two gregarine parasites, Protomagalhaensia granulosae and Gregarina cubensis (Apicomplexa: Eugregarinida) infecting the Death’s Head Cockroach, Blaberus discoidalis. Comparative Parasitology 70: 196-199. |
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Leander, B. S., R. E. Clopton, and P. J. Keeling. 2003. Phylogeny of gregarines (Apicomplexa) as inferred from small-subunit rDNA and beta-tubulin. International Journal of Systematic and Evolutionary Microbiology 55: 345-354. |
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Clopton, R. E., and C. M. Nolte. 2002. Clitellocephalus americanus n. gen., n. sp. (Apicomplexa: Eugregarinida: Gregarinidae) from Cratacanthus dubius (Coleoptera: Carabidae: Harpalinae) in the Nebraska Sandhills and Clitellocephalus ophoni n. comb. (Apicomplexa: Eugregarinida: Gregarinidae) from Ophonus pubescens (Coleoptera: Carabidae: Harpalinae) in Sète, France. Journal of Parasitology 88: 750-757. |
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Clopton, R. E., and A. Smith. 2002. Efficacy of oral sulfadimethoxine against two gregarine parasites, Protomagalhaensia granulosae and Gregarina cubensis (Apicomplexa: Eugregarinida) infecting the Death’s Head Cockroach, Blaberus discoidalis. Journal of Parasitology 88: 786-789. |
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Clopton, R. E. 2002. Phylum Apicomplexa Levine, 1970: Order Eugregarinorida Léger, 1900. Pages 205-288 in: Illustrated Guide to the Protozoa, 2nd edition, J. J. Lee, G. Leedale, D. Patterson, and P. C. Bradbury, eds. Society of Protozoologists, Lawrence, Kansas. |
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Cook, T. J., J. Janovy, Jr., and R. E. Clopton. 2001. Epimerite-host epithelium relationships among eugregarines parasitizing the damselflies Enallagma civile and Ischnura verticalis. Journal of Parasitology 87: 988-996. |
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Clopton, R. E. 2000. Stylocephalus occidentalis n. sp.(Apicomplexa: Eugregarinida: Stylocephalidae) from Trimytis pruinosa (Coleoptera: Tenebrionidae) in the Nebraska Sandhills. Journal of Parasitology 86: 560-565. |
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Clopton, R. E., and R. R. Kula. 1999. Amoebogregarina nigra n. gen., n. comb. (Apicomplexa: Gregarinidae) From Adult Melanoplus differentialis (Orthoptera: Acrididae) in Southeastern Nebraska. Journal of Parasitology 85: 321-325. |
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Clopton, R. E. 1999. Revision of the genus Xiphocephalus and description of Xiphocephalus ellisi n. sp.(Apicomplexa: Eugregarinida: Stylocephalidae) from Eleodes opacus (Coleoptera: Tenebrionidae) in the western Nebraska Sandhills. Journal of Parasitology 85:84-89. |
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Lucarotti, C. J., T. L. Leclerc, and Clopton, R. E. 1998. Incidence of Leidyana canadensis (Apicomplexa: Eugregarinida) in Lambdina fiscellaria fiscellaria larvae (Lepidoptoera: Geometridae). Canadian Entomologist 130: 583-594. |
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Clopton, R. E. 1998. Revision of the genus Torogregarina and description of Torogregarina sphinx n. Sp. (Apicomplexa: Eugregarinida) from a Missouri river bank beetle, Bembidion laevigatum (Coleoptera: Carabidae), in southeastern Nebraska. Journal of Parasitology 84: 823-827. |
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Clopton, R. E. and C. J. Lucarotti. 1997. Leidyana canadensis N. Sp. (Apicomplexa: Eugregarinida) from larval Eastern Hemlock Looper, Lambdina fiscellaria fiscellaria (Lepidoptoera: Geometridae). Journal of Eukaryotic Microbiology 44: 383-387. |
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Janovy, Jr., S. D. Snyder, and R. E. Clopton. 1997. Evolutionary constraints on population structure: the parasites of Fundulus zebrinus (Pisces: Cyrinodontidae) in the South Platte River of Nebraska. Journal of Parasitology 83: 584-592. |
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Clopton, R. E. 1995. Leidyana migrator n. sp. (Apicomplexa: Leidyanidae) from the Madagacar hissing cockroach, Gromphadorhina portentosa (Insecta: Blattodea). Invertebrate Biology 114: 271-278. |
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Clopton, R. E. and R. E. Gold. 1996. Host Specificity of Gregarina blattarum among Five Species of Domiciliary Cockroaches. Journal of Invertebrate Pathology 67: 219-223. |
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Clopton, R. E. 1995. Domadracunculus janovyii n. gen., n. sp., (Apicomplexa: Actinocephalidae) described from adults of Ischnura verticalis (Odonata: Zygoptera) in Texas. Journal of Parasitology 81:256--260. |
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Clopton, R. E. and R. E. Gold. 1995. Effects of pH on excystation in Gregarina cuneata and Gregarina polymorpha (Eugregarinida: Gregarinidae). Journal of Eukaryotic Microbiology 42: 540-544. |
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Percival, T. J., R. E. Clopton and J. Janovy, Jr. 1995. Hoplorhynchus acanthatholius N. Sp. and Steganorhynchus dunwoodii N. Gen. N. Sp. (Apicomplexa: Eugregarinorida) described from Coenagrionid Damselflies. Journal of Eukaryotic Microbiology 42: 406-410. |
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Janovy, Jr., R. E. Clopton, D. A. Clopton, S. Snyder, A. Efting and L. Krebs. 1994. Species density distributions as null models for ecologically significant interactions of parasite species in an assemblage. Ecological Modeling 77: 189-196. |
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Clopton, R. E. and R. E. Gold. 1993. Distribution and Seasonal and diurnal activity patterns of Eutrombicula alfreddugesi (Acari: Trombiculidae) in a forest edge ecosystem. Journal of Medical Entomology 30: 47--53. |
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Clopton, R. E. and J. Janovy, Jr. 1993 Developmental niche structure in the gregarine assemblage parasitizing Tenebrio molitor. Journal of Parasitology 79:701--709. |
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Clopton R. E., T. J. Percival and John Janovy, Jr. 1993. Nubenocephalus nebraskensis N. Gen., N. Sp. (Apicomplexa: Actinocephalidae) described from Argia bipunctulata (Odonata: Zygoptera). Journal of Parasitology 79: 533--537. |
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Ferdig, T., M. A. McDowell, R. E. Clopton and Janovy, J. Jr. 1993. Patterns of morphological variation of Salsuginus yutanensis (Monogenera: Ancyrocephalidae) over space and time. Journal of Parasitology 79: 744-750. |
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Clopton, R. E. and R. E. Gold. 1992. Bite-Count evaluation of the repellency of N,N-Diethyl-3-methylbenzamide to larval Eutrombicula alfreddugesi (Acari: Trombiculidae). Journal of Medical Entomology 29: 858-863. |
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Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. Host Stadium Specificity in the gregarine assemblage parasitizing Tenebrio molitor. Journal of Parasitology 78: 334--337. |
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Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. Gregarina coronata N. Sp. (Apicomplexa: Eugregarinida) described from adults of the Southern Corn Rootworm, Diabrotica undecimpunctata howardi. Journal of Protozoology 39: 417--420. |
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Janovy, Jr., R. E. Clopton and T. J. Percival. 1992. The Roles of ecological and evolutionary influences in providing structure to parasite species assemblages. Journal of Parasitology 78: 630-640. |
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Clopton R. E., T. J. Percival and John Janovy, Jr. 1991. Gregarina niphandrodes N. Sp. (Apicomplexa: Eugregarinorida) from adult Tenebrio molitor (L.) with oocyst descriptions of other gregarine parasites of the Yellow Mealworm. Journal of Protozoology 38:472--483. |
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Clopton R. E. 1991. A review of the Scaritinid Beetles (Coleoptera: Carabidae) of Nebraska. Transactions of the Nebraska Academy of Science 18: 53--65. |

The BASE URL for this site is http://science.peru.edu/gregarina © 2006 by Richard E. Clopton
Last Updated: Monday, January 02, 2006 © 2006 by Richard E. Clopton
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