R. E. Clopton

Richard E. Clopton, PhD

Professor of Biology
Editor, Comparative Parasitology
Downey Family Honors Chair in Science
Department of Natural Science
Peru State College
Peru, NE 68421
402/872-2284
402/872-2255
rclopton@oakmail.peru.edu

Education

1990-1993

PhD, University of Nebraska, Parasitology
Dissertation (under John Janovy, Jr.): "Specificity in the gregarine assemblage parasitizing Tenebrio molitor."

1987-1989

MS, University of Nebraska, Entomology
Thesis (under Roger E. Gold): "Some aspects of the biology and ecology of the common pest chigger mite, Eutrombicula alfreddugesi, Oudemans 1912."

1983-1987

BS, University of Nebraska, Agriculture, Entomology

Grantsmanship

1998

National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 1998-May 15, 1999; $10,000

1997

National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 1997-May 15, 1998; $10,000

Click on the PDF icon to download reprints in PDF format where available.

National Science Foundation.
Microbiological Survey & Inventory: Eugregarinida (Protista: Apicomplexa) parasitizing selected mandibulate arthropods of the Nebraska Sandhills. NSF DEB-9705179.
Funded: May 15, 1997-May 15, 2000; $199,780

Refereed Publications

Professor of Biology
Editor, Comparative Parasitology
Downey Family Endowed Honors Chair in Science
Department of Natural Science
Peru State College, Peru, NE
Lecture responsibilities (1995-Present)
Introductory Biology, Honors Biology, Zoology, Botany, Horticulture, Human Physiology, Genetics, Plant Systematics, Invertebrate Zoology, Entomology, Comparative Vertebrate Anatomy, Human Clinical Parasitology, and Experimental Biology
Scientific research
Systematics & Taxonomy of Gregarines parasitizing mandibulate arthropods; Evolutionary systematics; Evolution of symbiosis
Extramural funding by the National Science Foundation
Strong undergraduate research component
Summer Biological Survey & Inventory Expeditions, 1997 - 2005

1999

National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 1999-May 15, 2000; $10,000

2000

National Science Foundation:
Research Experience for Undergraduates
Supplemental grant to NSF DEB-9705179.
Funded: May 15, 2000-May 15, 2001; $10,000

Position

Richard E. Clopton, PhD

2004

National Science Foundation
Microbiological Survey & Inventory of gregarines parasitizing aquatic and riparian insects of the Texas Big Thicket.
NSF DEB 0340774
Funded: Oct 1. 2004-2007; $200,000

	Clopton, R. E. 2009. Phylogenetic Relationships, Evolution, and Systematic Revision of the Septate Gregarines (Apicomplexa: Eugregarinorida: Septatorina). Comparative Parasitology 76: 167-190.
	Clopton, R. E., T. J. Cook, and J. L. Cook. 2008. Gregarina tropica n. sp. (Apicomplexa: Eugregarinorida: Gregarinicae: Gregarinidae) Parasitizing the Brown-winged Earwig, Vostox brunneipennis (Dermaptera: Labiidae), in the Texas Big Thicket. Comparative Parasitology 75:215-227.
	Clopton, R. E., T. J. Cook, and J. L. Cook. 2008. Trichurispora wellgundis n. g., n. sp. (Apicomplexa: Eugregarinida: Hirmocystidae) parasitizing adult Water Scavenger Beetles, Tropisternus collaris (Coleoptera: Hydrophilidae) in the Texas Big Thicket. Comparative Parasitology 75: 82-91.
	Realzola, E., J. L. Cook, T. J. Cook, and R. E. Clopton. 2007. Composition of gyrinid aggregations in the East Texas Primitive Big Thicket. Coleopterist’s Bulletin 61:495-502.
	Hays, J. J., R. E. Clopton, T. J. Cook and J. L. Cook. 2007. Revision of the Genus Nubenocephalus and Description of Nubenocephalussecundus N. Sp. (Apicomplexa: Actinocephalidae) Parasitizing Adults of Argia sedula (Odonata: Zygoptera: Coenagrionidae) in the Primitive Texas Big Thicket, U.S.A. Comparative Parasitology. 74: 286-293.
	Clopton, R.E., T. J. Cook and J. L. Cook. 2007. Revision of GeneiorhynchusSchneider, 1875 (Apicomplexa: Eugregarinida: Actinocephalidae: Acanthosporinae) with Recognition of 4 New Species of Geneiorhynchus and Description of Geneiorhynchus manifestus N. Sp. Parasitizing Naiads of the Green Darner, Anax junius (Odonata: Aeshnidae) in the Texas Big Thicket. Comparative Parasitology 74: 273-285.
	Clopton, R. E., and J. J. Hays. 2006. Revision of the Genus Protomagalhaensia and Description of Protomagalhaensia wolfi n. comb. (Apicomplexa: Eugregarinida: Hirmocystidae) and Leidyana haasi n. comb. (Apicomplexa: Eugregarinida: Leidyanidae) Parasitizing the Lobster Cockroach, Nauphoeta cinerea (Dictyoptera: Blaberidae) Comparative Parasitology 73: 137-156.
	Clopton, R. E. 2006. Two New Species of Xiphocephaus in Eleodes tricostata and Eleodes fusiformis (Coleoptera: Tenebrionidae: Eleodini) from The Sandhills of Western Nebraska. Journal of Parasitology 92: 569-577.
	Welsh, D., R. E. Clopton, and L. Parris. 2006. Differential Temperature Acclimatization Responses in the Membrane Phospholipids of Posthodiplostomum minimum and its Second Intermediate Host, Lepomis macrochirus. Journal of Parasitology 92: 764-769.
	Hays, J., R. E. Clopton, D. L. Cappaert, and D. R. Smitley. 2004. Revision of the genus Stictospora and description of Stictospora villani, n. sp. (Apicomplexa: Eugregarinida: Actinocephalidae) from the Japanese beetle Popillia japonica (Coleoptera: Scarabaeidae) in Michigan, USA. Journal of Parasitology 90:1450-1456.
	Clopton, R. E. 2004. Standard nomenclature and metrics of plane shapes for use in gregarine taxonomy. Comparative Parasitology 71: 130-140.
	Clopton, R. E. 2004. Calyxocephalus karyopera g. nov., sp. nov. (Eugregarinorida: Actinocephalidae: Actinocephalinae) from the ebony jewelwing damselfly Calopteryx maculata (Zygoptera: Calopterygidae) in southeast Nebraska: implications for mechanical prey-vector stabilization of exogenous gregarine development. Comparative Parasitology 71: 141-153.
	Clopton, R. E., T. J. Cook, and J. L. Cook. 2004. Naiadocystis phykoterion n. gen., n. sp. (Apicomplexa: Eugregarinida: Hirmocystidae) from the mexican pygmy grasshopper Paratettix mexicanus (Orthoptera: Tetrigidae) in the Texas Big Thicket with recognition of three previously described species of Naiadocystis. Journal of Parasitology 90: 301-307.
	Smith A., and R. E. Clopton. 2003. Efficacy of oral metronidazole and potassium sorbate against two gregarine parasites, Protomagalhaensia granulosae and Gregarina cubensis (Apicomplexa: Eugregarinida) infecting the Death’s Head Cockroach, Blaberus discoidalis. Comparative Parasitology 70: 196-199.
	Leander, B. S., R. E. Clopton, and P. J. Keeling. 2003. Phylogeny of gregarines (Apicomplexa) as inferred from small-subunit rDNA and beta-tubulin. International Journal of Systematic and Evolutionary Microbiology 55: 345-354.
	Clopton, R. E., and C. M. Nolte. 2002. Clitellocephalus americanus n. gen., n. sp. (Apicomplexa: Eugregarinida: Gregarinidae) from Cratacanthus dubius (Coleoptera: Carabidae: Harpalinae) in the Nebraska Sandhills and Clitellocephalus ophoni n. comb. (Apicomplexa: Eugregarinida: Gregarinidae) from Ophonus pubescens (Coleoptera: Carabidae: Harpalinae) in Sète, France. Journal of Parasitology 88: 750-757.
	Clopton, R. E., and A. Smith. 2002. Efficacy of oral sulfadimethoxine against two gregarine parasites, Protomagalhaensia granulosae and Gregarina cubensis (Apicomplexa: Eugregarinida) infecting the Death’s Head Cockroach, Blaberus discoidalis. Journal of Parasitology 88: 786-789.
	Clopton, R. E. 2002. Phylum Apicomplexa Levine, 1970: Order Eugregarinorida Léger, 1900. Pages 205-288 in: Illustrated Guide to the Protozoa, 2^nd edition, J. J. Lee, G. Leedale, D. Patterson, and P. C. Bradbury, eds. Society of Protozoologists, Lawrence, Kansas.
	Cook, T. J., J. Janovy, Jr., and R. E. Clopton. 2001. Epimerite-host epithelium relationships among eugregarines parasitizing the damselflies Enallagma civile and Ischnura verticalis. Journal of Parasitology 87: 988-996.
	Clopton, R. E. 2000. Stylocephalus occidentalis n. sp.(Apicomplexa: Eugregarinida: Stylocephalidae) from Trimytis pruinosa (Coleoptera: Tenebrionidae) in the Nebraska Sandhills. Journal of Parasitology 86: 560-565.
	Clopton, R. E., and R. R. Kula. 1999. Amoebogregarina nigra n. gen., n. comb. (Apicomplexa: Gregarinidae) From Adult Melanoplus differentialis (Orthoptera: Acrididae) in Southeastern Nebraska. Journal of Parasitology 85: 321-325.
	Clopton, R. E. 1999. Revision of the genus Xiphocephalus and description of Xiphocephalus ellisi n. sp.(Apicomplexa: Eugregarinida: Stylocephalidae) from Eleodes opacus (Coleoptera: Tenebrionidae) in the western Nebraska Sandhills. Journal of Parasitology 85:84-89.
	Lucarotti, C. J., T. L. Leclerc, and Clopton, R. E. 1998. Incidence of Leidyana canadensis (Apicomplexa: Eugregarinida) in Lambdina fiscellaria fiscellaria larvae (Lepidoptoera: Geometridae). Canadian Entomologist 130: 583-594.
	Clopton, R. E. 1998. Revision of the genus Torogregarina and description of Torogregarina sphinx n. Sp. (Apicomplexa: Eugregarinida) from a Missouri river bank beetle, Bembidion laevigatum (Coleoptera: Carabidae), in southeastern Nebraska. Journal of Parasitology 84: 823-827.
	Clopton, R. E. and C. J. Lucarotti. 1997. Leidyana canadensis N. Sp. (Apicomplexa: Eugregarinida) from larval Eastern Hemlock Looper, Lambdina fiscellaria fiscellaria (Lepidoptoera: Geometridae). Journal of Eukaryotic Microbiology 44: 383-387.
	Janovy, Jr., S. D. Snyder, and R. E. Clopton. 1997. Evolutionary constraints on population structure: the parasites of Fundulus zebrinus (Pisces: Cyrinodontidae) in the South Platte River of Nebraska. Journal of Parasitology 83: 584-592.
	Clopton, R. E. 1995. Leidyana migrator n. sp. (Apicomplexa: Leidyanidae) from the Madagacar hissing cockroach, Gromphadorhina portentosa (Insecta: Blattodea). Invertebrate Biology 114: 271-278.
	Clopton, R. E. and R. E. Gold. 1996. Host Specificity of Gregarina blattarum among Five Species of Domiciliary Cockroaches. Journal of Invertebrate Pathology 67: 219-223.
	Clopton, R. E. 1995. Domadracunculus janovyii n. gen., n. sp., (Apicomplexa: Actinocephalidae) described from adults of Ischnura verticalis (Odonata: Zygoptera) in Texas. Journal of Parasitology 81:256--260.
	Clopton, R. E. and R. E. Gold. 1995. Effects of pH on excystation in Gregarina cuneata and Gregarina polymorpha (Eugregarinida: Gregarinidae). Journal of Eukaryotic Microbiology 42: 540-544.
	Percival, T. J., R. E. Clopton and J. Janovy, Jr. 1995. Hoplorhynchus acanthatholius N. Sp. and Steganorhynchus dunwoodii N. Gen. N. Sp. (Apicomplexa: Eugregarinorida) described from Coenagrionid Damselflies. Journal of Eukaryotic Microbiology 42: 406-410.
	Janovy, Jr., R. E. Clopton, D. A. Clopton, S. Snyder, A. Efting and L. Krebs. 1994. Species density distributions as null models for ecologically significant interactions of parasite species in an assemblage. Ecological Modeling 77: 189-196.
	Clopton, R. E. and R. E. Gold. 1993. Distribution and Seasonal and diurnal activity patterns of Eutrombicula alfreddugesi (Acari: Trombiculidae) in a forest edge ecosystem. Journal of Medical Entomology 30: 47--53.
	Clopton, R. E. and J. Janovy, Jr. 1993 Developmental niche structure in the gregarine assemblage parasitizing Tenebrio molitor. Journal of Parasitology 79:701--709.
	Clopton R. E., T. J. Percival and John Janovy, Jr. 1993. Nubenocephalus nebraskensis N. Gen., N. Sp. (Apicomplexa: Actinocephalidae) described from Argia bipunctulata (Odonata: Zygoptera). Journal of Parasitology 79: 533--537.
	Ferdig, T., M. A. McDowell, R. E. Clopton and Janovy, J. Jr. 1993. Patterns of morphological variation of Salsuginus yutanensis (Monogenera: Ancyrocephalidae) over space and time. Journal of Parasitology 79: 744-750.
	Clopton, R. E. and R. E. Gold. 1992. Bite-Count evaluation of the repellency of N,N-Diethyl-3-methylbenzamide to larval Eutrombicula alfreddugesi (Acari: Trombiculidae). Journal of Medical Entomology 29: 858-863.
	Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. Host Stadium Specificity in the gregarine assemblage parasitizing Tenebrio molitor. Journal of Parasitology 78: 334--337.
	Clopton R. E., T. J. Percival and John Janovy, Jr. 1992. Gregarina coronata N. Sp. (Apicomplexa: Eugregarinida) described from adults of the Southern Corn Rootworm, Diabrotica undecimpunctata howardi. Journal of Protozoology 39: 417--420.
	Janovy, Jr., R. E. Clopton and T. J. Percival. 1992. The Roles of ecological and evolutionary influences in providing structure to parasite species assemblages. Journal of Parasitology 78: 630-640.
	Clopton R. E., T. J. Percival and John Janovy, Jr. 1991. Gregarina niphandrodes N. Sp. (Apicomplexa: Eugregarinorida) from adult Tenebrio molitor (L.) with oocyst descriptions of other gregarine parasites of the Yellow Mealworm. Journal of Protozoology 38:472--483.
	Clopton R. E. 1991. A review of the Scaritinid Beetles (Coleoptera: Carabidae) of Nebraska. Transactions of the Nebraska Academy of Science 18: 53--65.

Dr. R. E. Clopton